Pros
A great collection of people. The IT department is staffed with extremely talented people who get along with each other very well. The work is interesting: modern tech stacks and (depending on the team) the opportunity to work in the fascinating genetic processing that forms the backbone of the company's product. Managers (below the Director level) really care about their employees and go to bat for them. It's possible to negotiate a strong starting salary.
Cons
There's a lack of clear vision for the department; projects are started and never finished, people are shuffled around to different projects. The C-level has a clear lack of interest in anything not directly related to the customer-facing Family Tree DNA product. The work environment is an extremely loud, overcrowded open office, and upper management is resistant to remote work. Do not accept a job with this company until you've seen where you'll be working. The CEO decided to allow the FBI access to its customers' genetic data without notifying the customers. They also kept it secret from the employees, who became implicit in violating user privacy without even knowing about it until an article was published in BuzzFeed. They've since decided to rebrand and use their work with the FBI in their marketing. This is not a company to spend years at and grow through the ranks. They're extremely stingy with raises, especially when a promotion or role change is involved. If you start out not paid well, you're never going to get raises up to the point where you are. Modest pay raises for promotions (for example, from customer service to QA or QA to developer) were delayed by months. There are three women in the entire IT department, and no women developers. The culture, set by example from the top, is not inclusive or particularly respectful of diverse points of view.